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The frontal sinuses are cavities inside the frontal bone located at the junction between the face and the cranial vault and close to the brain. Despite a long history of study, understanding of their origin and variation through evolution is limited. This work compares most hominin species’ holotypes and other key individuals with extant hominids. It provides a unique and valuable perspective of the variation in sinuses position, shape, and dimensions based on a simple and reproducible methodology. We also observed a covariation between the size and shape of the sinuses and the underlying frontal lobes in hominin species from at least the appearance of Homo erectus. Our results additionally undermine hypotheses stating that hominin frontal sinuses were directly affected by biomechanical constraints resulting from either chewing or adaptation to climate. Last, we demonstrate their substantial potential for discussions of the evolutionary relationships between hominin species. Variation in frontal sinus shape and dimensions has high potential for phylogenetic discussion when studying human evolution.

In recent years, research into the occurrence of bats at the Dutch North Sea has shown that there is regular seasonal migration over sea. However, so far, little is known about their migration ecology, the fatality risks at offshore wind turbines, and the number of individuals migrating over sea. Since the Dutch government wants to boost the further development of wind energy production in the southern North Sea, the Ministry of Economic Affairs commissioned to Rijkswaterstaat a Wind at Sea Ecological Programme (in Dutch: Wozep). This study, as part of the Wozep-project Behaviour and Collision Risk of Bats (Bats_2), investigates how telemetry can be applied to gain insight into migratory movements of bats over land and over sea and individual bat behaviour near and in offshore wind farms. To find out whether it is wise to continue and further develop telemetry research in the context of the Wozep programme, we first identified potential telemetry methods for small bats based on a desk study and selected the most promising method for the application of telemetry. Members of the team attended an international workshop on telemetry in Lund (Sweden) to gather practical technical knowledge, gain insight in data-management standards, and increase their international network. Several field tests were conducted to test the equipment and explore the possibilities of tracking. Finally, suitable locations for bat trapping in bat boxes and for the use of a Heligoland trap were identified. There are several options to track bats with radio telemetry during their migration at the coast and over sea. However, for long-term monitoring of multiple individuals, establishing a grid of stationary receivers is the only feasible option. Eight field tests were carried out to test the performance of the technical infrastructure. Each of the tests was unique and set up to test the signal strength/detection range for a certain type of antenna or a certain transmitter/receiver constellation. We showed that detections over at least 6 km are possible, and likely more than 10 km can be achieved. Precondition is that the receiving stations must be installed at high structures (lighthouses, buildings, masts) or hills, installing them a few meters off ground level will lead to detection ranges just over one km. Furthermore we explored the possibilities of calculating movement tracks with a setup of several receivers using different estimation methods. These experiments indicated that a relatively high accuracy (of c. 100 m) can be reached estimating the location when signals of different receiving stations are combined. It is likely that the accuracy can be improved by estimating the bearings of the received signals based on the signal strength of different antennas. Even further improvement seems possible by assessing the (likely) flight route with a state-space model. There are plenty of locations with bat boxes, especially in the province of Noord-Holland, where potentially hundreds of bats can be captured during migration stopovers, though it is not known how many of these are likely to be migrants. In addition, we identified four locations where actively migrating bats can potentially be captured with an Heligoland trap. In conclusion, we are confident that telemetry can be successfully applied to study migratory movements of bats over land and over sea and individual bat behaviour near and in offshore wind farms. Joining a wildlife tracking system like Motus (Canada) in order to enlarge the data collection, is highly recommended. Motus-members can get detections from both

Sub-Antarctic islands are expected to show a high degree of endemicity due to their remoteness. However, biogeographic affinities in the sub-Antarctic remain poorly understood, especially in the marine realm. Sub-Antarctic islands being at the crossroads between Antarctic and cold temperate regions, biodiversity characterization and biogeographic analyses are a priority for monitoring and rapidly assessing variations associated with environmental changes. One underexplored sub-Antarctic area is Crozet, a protected archipelago located halfway between Antarctica and South Africa. In this study, we investigated the shallow-water Crozet macrofaunal diversity, distribution patterns and biogeographic affinities based on the examination of fieldwork specimens via a thorough morphological identification and a genetic characterisation. The resulting dataset provides an important baseline for further studies and conservation strategies, compiling the first genetic and taxonomic database for the Crozet archipelago. In total, 100 morphotypes were found, belonging to nine different phyla, among whicharthropods(32), molluscs (18)and echinoderms (17) were the richest. Forty-seven morphotypes were identified to the species level, among which 20 were reported in Crozet for the first time. This confirms that Crozet is a poorly known region, even compared to other sub-Antarctic areas. A large proportion of species (62%) had circum Southern Ocean or circum sub Antarctic distributions. These species were mostly shared with Kerguelen (72%), the Magellan Province (64%), and Prince Edward Islands (64%), confirming the patterns found in macroalgae and specific macrofaunal groups. However, this large-distribution statement needs to be counterbalanced by the detection (genetic data) of more restricted distributions than expected in four study cases (the tanaid Apseudes spectabilis, the nudibranch Doris kerguelenensis, the polychaete Neanthes kerguelensis and the chiton Hemiarthrum setulosum). Considering that most morphotypes had no genetic data available from other regions, the proportion of morphotypes with restricted distribution is likely to increase alongside future investigations. In addition, we also found a few cases of unrecognized diversity that might lead to the descriptions of new species, some likely to be endemic to Crozet (e.g., within the polychaete genus Harmothoe and the bryozoan genus Antarctothoa). Altogether, this stresses the need to maintain conservation efforts in Crozet and pursue integrative investigations in order to highlight and protect its unusual diversity

RÉSUMÉ Les observations pétrographiques en lames-minces et les analyses au microscope électronique à balayage complètent les observations macroscopiques et mésoscopiques faites sur des tessons appartenant à deux espaces d’occupation rubanée situés à Velroux (Grâce-Hollogne, Province de liège, Belgique). Quatre groupes et plusieurs sous-groupes de pâtes céramiques ont été identifiés et caractérisés. Les sources locales et régionales des matières premières utilisées par les potiers rubanés sont discutées et ouvrent des perspectives sur les utilisations des matières premières sur la grande aire rubanée. MOTS-CLÉS : Rubané, pétrographie en lames-minces, microscope électronique à balayage, EDS, sources des matières premières, Hesbaye, Belgique. SUMMARY/ABSTRACT Petrographic observations in thin sections and scanning electron microscopic analyses complete the macroscopic and mesoscopic observations made on pottery sherds belonging to two occupation area belonging to the Linear Pottery culture and located at Velroux (Grâce-Hollogne, Province of Liege, Belgium). Four groups of ceramic pastes and some sub-groups were identified and characterized. Local and regional sources of raw materials used by the LBK potters are discussed and open perspectives on the uses of raw materials in the large Linear Pottery Culture area. KEYWORDS: Linear Pottery Culture, thin-section petrography, scanning electron microscope, EDS, raw material sourcing, Hesbaye, Belgium.

One of the greatest challenges of the last decades in the fight against climate change has been to achieve net-zero emissions by mid-century. According to the US EPA (2016), in 2014, global anthropogenic emissions of carbon dioxide (CO2) accounted for ~64% of the greenhouse effect. Carbon dioxide capture and storage (CCS) plays an irreplaceable part as a mitigation technology that avoids CO2 emissions at their source and bridges the transition into a non-carbon-based energy future. The International Energy Agency (IEA) estimates that the need to store CO2 will grow from 40 Mt/y at present to more than 5000 Mt/y by 2050. Additionally, in the IEA’s Sustainable Development Scenario, which aims for global net-zero CO2 emissions from the energy sector by 2070, CCS needs to become a global industry supporting emissions reductions across the overall energy system. CCS technologies essentially consist of capturing and compressing the CO2 at the source and then transport it towards deep suitable rock formations where it is injected to be permanently stored. The key to successful and permanent CO2 storage is the proper analysis and characterization of the reservoir and seal formation. Among the types of reservoir suitable for CO2 storage are unmined coal beds, depleted oil and gas fields, EOR/EGR, saline aquifers, man-made caverns, and basaltic formations (IPCC, 2005). The storage capacity of any of these reservoirs is the subsurface commodity whose quantities and properties are assessed when existing data is provided. Capacity estimations bring their own level of uncertainty and complexity according to the scale at which they are addressed and the nature of the geological conditions of the reservoir. This degree of uncertainty should be accounted for in every estimation (Bradshaw et al., 2007) Resource classification systems (RCS) are frameworks that establish the principles and boundaries for each level of capacity assessment. By making use of these frameworks, it is possible to properly allocate the stage of development of a resource (United Nations, 2020). For every level of assessment, the principles of the estimation change and so do the scale and purpose. As the analysis moves forward, a prospective site develops and exhaustive information is acquired, initial estimations are adjusted, and uncertainty is likely to reduce. Additionally, different economic, technical, regulatory, environmental and societal factors are integrated into the assessment to bring the estimations under present conditions. For instance, if the storage capacity is to be matched with a CO2 source, detailed simulations and analyses regarding injectivity, supply rate, potential routes and economic distances must be performed to achieve a realistic estimation. However, an assessment where the main goal is to merely quantify the space available to store CO2 in a reservoir, does not consider the aforementioned limitations and will carry higher risk and uncertainty in its estimation (Bradshaw et al., 2007). Even though resource classification systems provide a solid foundation for CCS projects, they do not provide the input parameters and analyses needed to reach every level of assessment. This is why storage capacity estimation methodologies go hand in hand with RCS given that the former can give information related to the parameters and constraints considered in the estimation. No standard process has been proposed that can be followed from the starting level of a CO2 storage capacity assessment until a fully developed carbon storage resource; that is, a CO2 storage site ready to become fully operational. This paper aims to develop a methodology where the fundamental steps needed to go through every level of the resource classification systems are standardized. This methodology intends to serve as a general baseline that, regardless of the geological settings and techno-socio-economic conditions, can be adopted for any CCS assessment. The proposed methodology is built by reviewing the available capacity estimation methods for every level of assessment and identifying social, technical and economic aspects that come into play as the resource is being developed. Considering that capacity estimation methodologies can vary their approach even for the same level of assessment, the rationales behind them are expected to be determined. Such rationales can be related to in-place policy restrictions, geographical economic behavior, or the nature of the parameters contemplated. Additionally, PSS, an in-house developed tool that can assess CO2 storage reservoirs at different levels, will be proposed within the methodology. This tool is a bottom-up geotechnical and economic forecasting simulator that can generate source-sink matching for CCS projects, where technical, economic, and geological uncertainties are handled through a Monte Carlo approach for limited foresight (Welkenhuysen et al., 2016). Acknowledgements This research is carried out under the LEILAC2 project, which receives funding from the European Union’s Horizon 2020 research and innovation program under grant agreement number 884170. The LEILAC2 consortium consists of: Calix Europe SARL, HeidelbergCement AG, Ingenieurbüro Kühlerbau Neustad GmbH (IKN), Centre for Research and Technology Hellas (CERTH), Bundesanstalt für Geowissenschaften und Rohstoffe (BGR), Politecnico di Milano (POLIMI), Geological Survey of Belgium (RBINS-GSB), ENGIE Laborelec, Port of Rotterdam, Calix Limited, CIMPOR-Indústria de Cimentos SA and Lhoist Recherche et Development SA. References Bradshaw, J., Bachu, S., Bonijoly, D., Burruss, R., Holloway, S., Christensen, N. P., & Mathiassen, O. M. (2007). CO2 storage capacity estimation: Issues and development of standards. International Journal of Greenhouse Gas Control, 1(1), 62–68. https://doi.org/10.1016/S1750-5836(07)00027-8 IPCC. (2005). Carbon Dioxide Capture and Storage. https://www.ipcc.ch/report/carbon-dioxide-capture-and-storage/ United Nations. (2020). United Nations Framework Classification for Resources: Update 2019. UN. https://doi.org/10.18356/44105e2b-en US EPA. (2016). Global Greenhouse Gas Emissions Data. US EPA. https://www.epa.gov/ghgemissions/global-greenhouse-gas-emissions-data Welkenhuysen, K., Brüstle, A.-K., Bottig, M., Ramírez, A., Swennen, R., & Piessens, K. (2016). A techno-economic approach for capacity assessment and ranking of potential options for geological storage of CO2 in Austria. Geologica Belgica. http://dx.doi.org/10.20341/gb.2016.012

Knowledge on the environmental impacts of geothermal energy is of major importance to understand the role this technology could play in the transition towards sustainable energy systems. Life cycle analysis (LCA) methodology is a widely used tool for assessing the environmental impacts of products and systems, which has been implemented numerous times on geothermal systems. Previous reviews on geothermal LCA studies identify large variability on the reported environmental impacts. In this work we aim to provide a more in-depth analysis to explain the variability across the different LCAs. We review 28 LCA studies on geothermal energy published between 2005 and 2020, following a four step reviewing sequence; in step 1 we identify the LCA methodological choices and the plant geo-technical characteristics, in step 2 we identify the LCA results and the LCI inputs, in step 3 we perform contribution analysis based on the reported results and in step 4 we investigate the sensitivity and scenario analysis performed in the studies. If the data is available we triangularly evaluate the reported impacts considering a) the plants’ geo-technical characteristics, b) the hotspot analyses results and c) the Life cycle inventory (LCI) inputs. We focus our analysis on the six most frequently assessed impact indicators (GWP, AP, HTP, FETP, CED, ADP)* and distinguish between the different energy conversion technologies used for geothermal energy exploitation. This way we aim to provide a more transparent picture on the variability of environmental impacts across the LCAs by focusing on the environmental hotspots and on the cause-effect relationships between geo-technical parameters and the environmental impacts. We also aim for drawing LCA guidelines for future LCA studies on geothermal systems and proposing methods for impact mitigation. The variability on the LCA results is caused by differences on the choices of the LCA practitioners, on the energy conversion technologies used, on geological parameters and on plant design parameters. Most studies focus on the GWP and AP impacts, while information for the rest of the impacts is much more limited. For flash and dry steam power plants the direct emissions of non-condensable gases (NCGs) emerging can cause high GWP, AP, FETP and HTP impacts depending on the geofluid’s composition. The CED and ADP impacts are dominated by the steel and diesel consumption during the development of the wells. Thus differences on the geo-technical parameters determining the power output and the total material and energy consumption cause the variability on the reported results. Direct emissions of NCGs do not emerge in plants utilizing binary technology. In these plants the development of the wells dominates the impacts and this phenomenon is more intense when EGS-binary plants are investigated due to the large depth drilled. Also the production of the working fluid used in the ORC and its annual leakage can highly affect the GWP impact in these plants depending on the type of working fluid used. In heating plants high amounts of grid-electricity are needed for the plant operation as no power is produced. Therefore differences in the fossil-fuel-intensity of the electricity mix supplying the plant can result in large variability. The choice of the LCA practitioner to include or not the heat distribution network in the boundaries of the system also affects the results, while a significant portion of the impacts is caused during the development of the wells. Combined heat and power plants using flash or binary technology present similar results. However the co-production of heat and power is expected to lead to some benefits. A direct correlation between the GHGs and the NH3/H2S direct emissions with the GWP and AC impacts, respectively, is observed for flash and dry steam power plants. Direct emissions are determined by the geofluid composition which highly varies between different reservoirs. For mitigating these impacts the installation of abatement systems shall be considered, while the identification of the geofluid composition and of the natural emissions emerging prior to the plant development is suggested for estimating the actual anthropogenic emissions. For plants utilizing binary technology and heating plants it is observed that higher capacity generally leads to lower GWP and AP impacts per functional unit. The capacity is a product function of the temperature and production flow. Similar observation can be extracted for the temperature while this is not the case for the flow. No clear correlation can be seen between the impacts and the depth. This is because larger depths lead –on the one hand– to higher impacts because of higher material and energy consumption which are compensated –on the other hand– to the increase on the fluid temperature and flow. For mitigating impacts caused during the construction phase the use of renewable energy sources for supplying the machinery used is suggested, while proper fluid re-injection should be designed for keeping the capacity constant during the operation. Also for binary plants the working fluid shall be selected such that its GWP impact is low, while for heating plants the installation of a small ORC unit shall be considered if the conditions are appropriate for meeting the pumping needs of the plant. The reviewed studies show that geothermal energy exploitation can lead to significant environmental benefits compared to fossil sources, as most of the times the impacts caused by geothermal plants are in the range of other renewable sources. Further research is needed on deep geothermal energy exploitation to better understand its environmental impacts. A significant portion of the impacts is caused during the operation of the plants regardless of the technology used (direct emissions, electricity consumption, working fluid losses, make-up well drilling). All of the LCA studies reviewed are static LCAs. Thus a dynamic LCA framework considering the time aspect is needed for better estimations of the environmental impacts. Also consequential LCAs on geothermal energy plants need to be conducted in order to assess how the global environmental impacts may change by the wider implementation of geothermal energy. In addition, future LCA studies shall also focus on environmental impacts other than the GWP as information regarding them is limited. Finally the sustainability of geothermal investments is to be further explored by investigating the social impacts of geothermal development and comparing them to other energy sources but also the financial aspect of such investments. Acknowledgments This research is carried out under the DESIGNATE project, which receives funding from the BELSPO BRAIN-be 2.0 research program under contract nr B2/191/P1/DESIGNATE. * GWP: Global Warming Potential, AP: Acidification Potential, HTP: Human Toxicity Potential, FETP: Freshwater EcoToxicity Potential, CED: Cumulative Energy Demand, ADP: Abiotic resources Depletion Potential