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Phytoplankton enhances the flocculation of suspended particulate matter in a tidal, turbid estuary
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RBINS Staff Publications 2025 OA
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Advancing Geothermal Potential in Wallonia (Belgium): Insights from Seismic Investigations of Dinantian Carbonates
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In response to climate and environmental challenges, the Walloon government is committed to achieving carbon neutrality by 2050 at the latest (including a 95% reduction in greenhouse gas emissions compared to 1990 levels). This goal is based on a progressive emissions reduction trajectory, with an interim target of a 55% reduction in greenhouse gas emissions by 2030 compared to 1990. To achieve its energy goals, Wallonia will need to adopt geothermal energy on a broad scale and initiate the development of this renewable sector within the region. Currently, geothermal energy represents only a small fraction of the energy mix, even though resources are available in the Sambre-Meuse valley for deep geothermal applications and across the entire region for shallow geothermal systems. Its use aligns fully with the energy transition currently underway in Wallonia. To assess the geothermal potential in Wallonia, gathering primary subsurface data remains essential. A pivotal part of this process involved seismic data acquisition under the DGE-ROLLOUT project (Interreg North-West Europe), completed in October 2023. This initiative generated several new cross-border seismic profiles, which have substantially reduced geological uncertainties, particularly along the BE/GE and BE/NL borders. These data supported the integration of mapped areas of the primary geothermal reservoir target, the Dinantian formation, and fostered knowledge sharing about potential geothermal aquifers. In Wallonia, the Geological Survey of Belgium (GSB) conducted 63.5 km of seismic surveys across Namur's eastern and western regions in December 2022 called GEOCOND 22, focusing on the extent of Dinantian limestones beneath the Midi-Eifelian Fault . Findings from this survey shed light on the geometry of the fault and the potential for Dinantian limestone occurrences at depths of 4-5 km, as well as Givetian-Frasnian carbonates at shallower depths (2-3 km) in the Condroz region. Additionally, reprocessing and reinterpretation of the Dekorp 1A line indicated a strong potential for Dinantian carbonates at depths suitable for deep geothermal exploration in Eastern Belgium Building on these promising results, the Walloon government has approvedurther large geophysical investigations in 2024. The WALSCAN project will gather approximately 400 km of seismic lines across three key areas: Charleroi, Liège, and Verviers/Eupen. Coordinated by GSB, Walscan brings together experienced partners like UMONS, ULG, and EPI Ltd, who contribute extensive expertise in geophysics, geothermal energy and regional geology. Scheduled for 2026, the seismic campaign will concentrate on identifying Dinantian carbonates at varying depths (1.5 km to 4 km) within densely urbanized areas, which align with regions of high energy demand.
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Geothermal Enery Use, country update for Belgium, EGC2025
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Geothermal energy is progressively gaining ground in Belgium, with tailored strategies emerging across its three Regions. Wallonia has undertaken a comprehensive modernization of its regulatory instruments, set ambitious renewable heat targets, and initiated large-scale subsurface exploration. Flanders is reinforcing its leadership in deep geothermal by targeting new geological formations, while improving shallow geothermal integration and subsurface governance. In the Brussels-Capital Region, efforts focus on incorporating shallow geothermal into urban energy planning through spatial zoning, technical potential mapping, and system monitoring. A suite of regional and European research projects (e.g. GEOCAMB, DESIGNATE, MORE-GEO, URGENT) have played a pivotal role in de-risking geothermal development by providing interdisciplinary tools that address geological complexity, economic feasibility, and environmental performance. Nevertheless, geothermal energy accounted for only 3.3% of Belgium’s renewable heat production in 2023, highlighting the need for accelerated deployment - especially in deep systems. Achieving carbon neutrality by 2050 will require stronger political commitment, harmonized regulatory frameworks, and targeted financial incentives. Ongoing pilot projects and scientific advances confirm geothermal energy's potential to become a cornerstone of Belgium’s sustainable heating transition.
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Do diversification and dispersals of early Eocene perissodactyls differ from those of other modern mammals?
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Perissodactyls make their first appearance in the fossil record during the Paleocene-Eocene Thermal Maximum (PETM), 56 Ma ago, at the same time as most modern placental mammal orders. However, their early diversification remains unclear since all main groups (Equidae, Isectolophidae, Ceratomorpha, Ancylopoda and Brontotheriidae) appear at the same time with numerous genera and are already diverse, contrary to other modern mammal orders, which have a much lower diversity. Moreover, several early perissodactyl genera seem endemic to each of the three Holarctic continents, rather than cosmopolitan as is the case for other mammal groups. We investigate the early evolution of perissodactyls by analyzing a new dataset focusing on early species to obtain a new expansive phylogeny. This large-scale phylogeny shows that the number of early perissodactyl genera was over-estimated and supports the synonymy of several genera. In addition, many species that were named as “Hyracotherium” in the past (or sometimes “Eohippus”, the “dawn-horse”) are not actually closely related to horses, including Hyracotherium itself, but can be considered as basal perissodactyls. Among the latter, at least Pliolophus and Cardiolophus experienced vast and fast dispersals between North America, Europe and Asia, during the PETM. This paleobiogeographic scenario much resembles the one of other earliest Eocene mammals, which comprise genera that are often found in at least two continents, such as in artiodactyls, primates, carnivorans, hyaenodontids, hyopsodontids, phenacodontids or rodents. Here, we show that the paleobiogeographic distribution of earliest perissodactyls does not differ from that of other modern mammals.
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Tomography of three articulated perissodactyl skeletons from Messel
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The perissodactyls from the UNESCO World Heritage site of Messel (MP11; around 47 Ma) are among the most well-known fossils from this site, and belong to at least five different genera: three equoids (Propalaeotherium, Eurohippus, and Hallensia) and two tapiromorphs (Lophiodon and Hyrachyus). Here, we investigated by X-ray microtomography at the DTHE (MateIS Laboratory, Lyon) the anatomy of three articulated skeletons from the Royal Belgian Institute of Natural Sciences collection: two specimens of Eurohippus messelensis, as well as the only accessible skeleton of Hallensia matthesi. The segmented 3D models allow us to visualize for the first time some cranial, dental and postcranial features that were previously concealed. Surprisingly, the tooth rows of the two specimens of E. messelensis show two different morphologies. One is typical of E. messelensis whereas the other shows all the diagnostic characters of E. parvulus expressed by Franzen (2006). However, based on the monograph on equoids from Messel by Franzen (2007), E. messelensis was only recognized in Messel, whereas E. parvulus was known from several other localities, notably in the Geiseltal, but not in Messel. We thus suggest that E. parvulus may have also been present in Messel, which raises the question of their potential synonymy. Finally, the preservation of the skeleton of Hallensia matthesi does not permit to observe the fine anatomy of the skeleton, but still permits to discuss important characters such as the number of fingers.
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RBINS Staff Publications 2025 OA
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The biodiversity of the Eocene Messel vertebrates based on the Belgian Collections
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The paleontological collections of the Royal Belgian Institute of Natural Sciences include a beautiful collection of 393 vertebrate specimens from the Messel Pit: 58 fish, 18 amphibians, 79 reptiles, 108 birds and 130 mammals. This collection is the largest Messel collection outside Germany and belongs to the „big four” (Smith et al. 2024). It results from a fieldwork partnership with the Seckenberg Research Institute, Frankfurt between 1982 and 1988, so a few years before the Messel pit was listed in 1995 as UNESCO World Heritage site. With a few exceptions, most of the specimens have been collected by the Belgians. However, most of the specimens have been prepared by German preparators. Here we show the Messel biodiversity based on the Brussels collection. The collection is relatively well diversified and contains 24 type and figured specimens. Among them are remarkable specimens such as the holotype of the hyaenodont carnivorous mammal Lesmesodon edingeri, the paratypes of the trogon bird Masillatrogon pumilio and embalonurid bat Tachypteron franzeni, the exquisitely 3-D prepared turtle lovers Allaeochelys crassesculpta and armored crocodylia Diplocynodon deponiae, and the only skeleton of the basal perissodactyl Hallensia matthesi housed in a public collection. The archives related to this collection are currently being digitized and several new studies are now based on micro-CT scan technology allowing to „discover” hidden characters.
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RBINS Staff Publications 2025 OA
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First crocodyliform fossils from the Eocene of Indonesia
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The Paleogene crocodyliform record is rich in taxonomic diversity, preserving crown and stemeusuchians alongside late-surviving neosuchians and notosuchians. This richness is mainly known from the extratropical latitudes of the New World and Eurasia, contrasting a poorly sampled tropical record. Within the tropics, the Paleogene of low-latitude southeast Asia is comparatively undersampled, with the Eocene Krabi Formation of Thailand representing the southernmost fauna. This crocodyliform fauna is composed of orientalosuchins and gavialoids, consistent with more northern records from southern Asia. Here we describe the first crocodyliform fossils from the Eocene of Indonesia, collected from the middle-late Eocene Talawi Member of the Sawahlunto Formation, west Sumatra. The Talawi Member represents a coastal swamp environment, preserving invertebrates, fish, and turtles along with rare frog, snake, and mammal fossils. Recovered crocodyliform fossils are often isolated and disarticulated, consisting of cranial remains including a partial braincase and jaw fragments, as well as relatively abundant teeth and osteoderms, vertebrae, and at least two autopodial elements. Procoelous vertebrae indicate eusuchian affinities. Osteoderms are diverse, with keeled and keelless squared and subrectangular forms. High and low-crowned fluted tooth morphotypes compare favorably to coeval gavialoids and orientalosuchins. A ziphodont form is present, potentially a planocraniid or sebecosuchian. External morphology of the braincase, such as narrow basioccipital tubera, a basioccipital plate bearing a crest, and a large basisphenoid exposure suggest a non-gavialoid identity. This is supported by preliminary observations of highresolution microCT scans, which imply a lateral carotid foramen ventral to the metotic foramen and laterally aligned medial and lateral eustachian foramina. The Sawahlunto fauna extends the southernmost geographical range of the Eocene southeast Asian crocodyliform fauna and represents the first multitaxon crocodyliform fauna from equatorial Asia. The ziphodont teeth either represent the last-occurring Asian planocraniids in a tropical refugium, or an incursion of Gondwanan sebecosuchians from India into the Asian tropics.
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Recent data on the enigmatic family Raoellidae: how aquatic…how cetacean?
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At the beginning of the 21st century, the discovery of an involucrum on the auditory bulla of Indohyus, a small raoellid artiodactyl from the Indian subcontinent, triggered the phylogenetic gathering of Raoellidae with the Cetacea clade, making them pivotal in addressing questions surrounding the land-to-water transition in Artiodactyla. Micro CT scan investigations and newly collected material from the Kalakot area (Jammu and Kashmir, India), recently greatly increased our knowledge of the cranial and dental features of this group. Here we present new data on cranial morphology of Raoellidae deriving from micro CT scan investigations of Indohyus indirae and Khirtharia inflata. This new body of data comprises investigations of the endocast of the braincase, the complete middle ear including the ossicles and auditory bulla, the petrosal bone, and the cast of the bony labyrinth. It also includes the reconstruction of the complete anterior dentition. This study gives us a comprehensive picture of the features of the Raoellidae head and improves our understanding of their semiaquatic habits. This increased knowledge of raoellids further extends the list of features they share with the first archeocetes, calling into question the definition of the clade Cetacea.
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RBINS Staff Publications 2025 OA
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Revision of the longirostrine crocodilian Thoracosaurus isorhynchus from the Maastrichtian–Danian of northwestern Europe
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“Thoracosaurs” are a group of longirostrine eusuchians known from Upper Cretaceous–lower Paleogene coastal deposits of Europe and North America. Their phylogenetic position is highly debated and varies from one study to the next, ranging from inclusion within Gavialoidea—implying ghost lineages of up to 100 million years—to exclusion from Crocodylia altogether. Despite these issues, many “thoracosaur” taxa have been understudied, suffering from incomplete or outdated descriptions and questionable taxonomic assignment. We present a comprehensive revision of Thoracosaurus isorhynchus, the most common European “thoracosaur.” Here, we redescribe the lectotype and paralectotypes of the species from the Maastrichtian of Mont-Aimé, France, and a skull from the Maastricht type area, The Netherlands. We also add new, previously undescribed cranial and extensive postcranial material from Mont-Aimé. Moreover, comparison with the type material of Thoracosaurus “scanicus” of the Danian of southern Sweden indicates that the latter is a junior synonym of T. isorhynchus, confirming the persistence of this species across the Cretaceous–Paleogene boundary. An array of phylogenetic analyses points to a close relationship between T. isorhynchus and the Cenomanian Portugalosuchus azenhae. This implicates an early emergence of “thoracosaurs” within Gavialoidea, highlighting the stratigraphic inconsistency of this group and a need for up to date descriptions of species belonging to this group.
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RBINS Staff Publications 2025 OA
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Building a Lithotectonic Framework for Belgium
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Almost all geological subdisciplines depend, to varying extents, on regional geological knowledge. Stratigraphic terminology is typically well-defined, while other concepts rely on generally accepted definitions or hierarchical schemes, such as palaeontological, structural and magmatic terminologies. This is much less the case for the regional geological building blocks. Their nomenclature is usually composed of a reference to a geographical locality and a geological term. Examples from Belgium include the (Anglo-)Brabant Massif, Campine Basin, and Malmedy Graben. Despite wide recognition, such terms often lack precise definitions and may even present conflicting interpretations across different contexts and authors. Even when their meanings have drifted or become less precise, these terms continue to be utilized. Increased awareness has led to significant yet isolated initiatives aimed at improving the structure and definition of regional geological information [1-3], recently brought together through pan-European cooperation [4]. Lithotectonic unit appears to be the most effective concept for encompassing all geological features. A lithotectonic unit is characterized by its composition, structural elements, mutual relations, and/or geological history [5]. Following a geotemporal conceptual approach, lithotectonic units are defined and bounded by relative limits in time and space [6]. Lithotectonic limits are planar features corresponding to geological events which have formed and define these units. Examples of lithotectonic units include orogens, terranes, sedimentary basins, and grabens, while examples of lithotectonic limits include deformation fronts, faults, and unconformities. This approach facilitates the organization and formalization of relationships between units and limits through ontologies. The data model can be linked to established ontologies, such as the ICS Geological Time Scale Ontology [7], and allows future extensions, such as attribution to orogenic cycles [2]. The associated concepts can be linked to 2D and 3D visualizations, thereby adding an important layer of knowledge to geological maps and models. Primary objective of the newly established Lithotectonic Working Group, under the National Commission for Stratigraphy in Belgium, is to create a comprehensive lithotectonic framework, that systematically defines and describes the main geological units and limits of Belgium. This initiative aligns closely with emerging standards currently being developed and implemented at European level [4] and largely based on GeoSciML [8]. [1] Hintersberger et al. 2017, Jb Geol B-A 157:195-207. [2] Németh 2021, Miner Slovaca 2:81-90. [3] Le Bayon et al. 2022: https://doi.org/10.1051/bsgf/2022017. [4] GSEU 2022-2027: https://doi.org/10.3030/101075609. [5] INSPIRE 2015: https://inspire.ec.europa.eu/theme/ge. [6] Piessens et al. 2024: https://doi.org/10.31223/X5RT28. [7] Cox & Richard 2005: https://doi.org/10.1130/GES00022.1. [8] GeoSciML 2016: http://www.opengis.net/doc/geosciml/4.1.
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