Easter Island (or Rapa Nui), internationally renowned for its megalithic statues, is the most isolated inhabited island of the Pacific. Archaeological surveys undertaken fromthe end of the 19th century led to the discovery of the remains of several hundred human individuals. The majority were buried in monuments (funerary stone platform called ahu) or in caves. This paper presents a study of the ancient Easter Islanders' diet through carbon and nitrogen stable isotope analysis of human tooth and bone collagen and, more particularly, evaluates the impact of gender, age, social status and location of burials. The 125 studied individuals are from 16 sites, which date mainly fromthe 17th to the 19th centuries. This anthropological material is housed at the Royal Belgian Institute of Natural sciences and the Father Sebastián Englert Anthropological Museum of Easter Island. One hundred and seven individuals showed well-preserved collagen. The stable isotope data provide new information on ancient Easter Islander dietary habits. They demonstrate gender disparity in access to food resources and show that children were breastfed until 3 years of age. Furthermore, the isotopic signatures cluster according to the place of burial (ahu) indicating family dietary specificities. Finally, our study reveals influences of social status on food intake: individuals from Ahu Nau Nau, which is said to be the royal ahu, display the highest nitrogen and carbon isotope values. A greater consumption of marine products may explain this distinction.
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The lower Eocene (Ypresian) Cambay Formation at Vastan Lignite Mine in Gujarat, western India, has yielded a rich vertebrate assemblage including the earliest modern mammals and oldest birds of the Indian subcontinent. Among the herpetological faunas, snakes, lizards and amphibians are abundant, but, strangely, lizards are only represented by agamids. Here we describe the agamid assemblage based on numerous, diverse and well-preserved dentaries, premaxillaries, and maxillaries. At least four taxa are present at Vastan. Vastanagama susanae is characterized by dentaries with a large symphyseal facet, three anterior pleurodont teeth followed by acrodont teeth presenting a main cusp bordered by two lateral crests; the teeth increase in size posteriorly toward the coronoid process. Tinosaurus indicus exhibits a subdental ridge between the tooth row and the Meckelian canal, pleurodont symphyseal teeth including one that can be caniniform, and acrodont and tricuspid posterior teeth with poorly differentiated lateral cusps. Two other taxa represent two new genera and species. The first taxon presents multicuspid acrodont teeth with the main cusp surrounded by two or three progressively smaller lateral cusps. The second taxon presents pleurodont anterior teeth followed by a few acrodont teeth and ending with three or four subacrodont teeth near the coronoid process. Our results confirm that Agamidae (assigned to the Acrodonta) is the only lizard group present at Vastan, whereas many other groups are already present in the Early Eocene on the other continents. Agamidae is considered to have had a Gondwanan origin, with 52 genera and 420 species of extant agamids known from Asia, Australia, Africa and a few from Southern Europe. The oldest occurrence of formally recognized Acrodonta is found in the Jurassic of India. Other fossil agamids are known in the Upper Paleocene of Kazakhstan, Paleocene and Eocene of China, Early Eocene of Europe, Eocene of North America, and Middle Eocene of Pakistan. The diversity of the agamids in India and the absence of other lizard groups at Vastan tentatively support the Out-of-India hypothesis for agamids.
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