Hervé Bocherens, Gennady Baryshnikov, and Wim Van Neer (2014)
Were bears or lions involved in salmon accumulation in the Middle Palaeolithic of the Caucasus? An isotopic investigation in Kudaro 3 cave
Quaternary International, 339-340:112-118.
Bone fragments of large anadromous salmon in the Middle Palaeolithic archaeological layers of Kudaro 3
cave (Caucasus) suggested fish consumption by archaic Hominins, such as Neandertals. However, large
carnivores such as Asiatic cave bears (Ursus kudarensis) and cave lions (Panthera spelaea) were also found
in the cave and could have been responsible for such an accumulation. The diet of these carnivores was
evaluated using carbon, nitrogen and sulphur isotopes in faunal bone collagen. The results suggest that
anadromous fish were neither part of the diet of either cave bear (vegetarian) or cave lion (predators of
herbivores from arid areas) and therefore provide indirect support to the idea that Middle Palaeolithic
Hominins, probably Neandertals, were able to consume fish when it was available.
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