A new basal raoellid artiodactyl (Mammalia) from the middle Eocene Subathu Group of Rajouri District, Jammu and Kashmir, northwest Himalaya, India

A new artiodactyl of moderate size is described on the basis of several dentaries and maxillae from the middle Eocene Subathu Group of the Kalakot area, Rajouri District, Jammu and Kashmir, India. Despite its general resemblance with the family Dichobunidae this taxon shares with Raoellidae two unambiguous characters: the presence of a hypoconulid on p4, and an asymmetrical P4. The position of the new taxon within the Cetacea / Raoellidae clade is strongly supported by eight non ambiguous synapomorphies, among which a cristid obliqua on lower molars anteriorly pointing towards the postectoprotocristid, and a P3 with only two roots. The new taxon is characterised by the following characters: a long symphysis; p3 and p4 with small parastylid and metastylid but no metaconid; lower molars with metaconid as the highest cusp, voluminous hypoconid, and absence of ‘hypolophid’; m1 and m2 with small paraconid, basally fused with metaconid, and small bifid tubercle-like hypoconulid; m3 with a crestiform paraconid; P3 and P4 with small protocone and shelf-like cingulae; upper molars with small paraconule and large metaconule (pseudohypocone); M1 and M2 with conical brachydont cusps; M3 with bunodont bulbous cusps; cristae and cingulae distinct and thick, presence of an ectoloph. The presence of a new primitive raoellid in the middle Eocene Subathu Group sheds new light on the phylogeny and paleobiogeography of basal raoellid artiodactyls. This work is partly funded by project BR/121/A3/PALEURAFRICA from the Belgian Science Policy Office.

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Paleontology