Nyungwe National Park (NP) is a mountainous region situated in the southwestern part of Rwanda on Congo-Nile watershed. In spite of the high biodiversity in pri- mates, birds and plants, no fish were reported to occur in the park, probably because of the cold temperatures of the rivers. An expedition in 2022 examined the fish diver- sity within the Nyungwe NP and its buffer zones. Additional sampling was performed in the main river draining the park into Lake Kivu: the Kamiranzovu. Three hundred and twenty specimens belonging to 13 species were collected. Specimens were col- lected only in the western part of the park, draining towards the Congo basin. The diversity within the park proper was limited to two putative species within the com- plex of Amphilius cf. kivuensis, which were caught on either side of the Kivu–Rusizi watershed. In contrast, a higher fish diversity, including one clariid species and two species of Enteromius, was observed in the rivers at a lower altitude of the buffer zone. However, the highest species diversity was found near the mouth of Kamiran- zovu River, including 11 species, of which 4 were non-native: the guppy Poecilia reti- culata, Astatotilapia burtoni, the blue-spotted tilapia Oreochromis leucosticus and the Egyptian mouth-brooder Pseudocrenilabrus multicolor.
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RBINS Staff Publications 2025
The anatomy of Cambaytherium, a primitive, perissodactyl-like mammal from the lower Eocene Cambay Shale Formation of Gujarat, India, is described in detail on the basis of more than 350 specimens that represent almost the entire dentition and the skeleton. Cambaytherium combines plesiomorphic traits typical of archaic ungulates such as phenacodontids with derived traits characteristic of early perissodactyls. Cambaytherium was a subcursorial animal better adapted for running than phenacodontids but less specialized than early perissodactyls. The cheek teeth are bunodont with large upper molar conules, not lophodont as in early perissodactyls; like perissodactyls, however, the lower molars have twinned metaconids and m3 has an extended hypoconulid lobe. A steep wear gradient with heavy wear in the middle of the tooth row suggests an abrasive herbivorous diet. Three species of Cambaytherium are recognized: C. thewissi (∼23 kg), C. gracilis (∼10 kg), and C. marinus (∼99 kg). Body masses were estimated from tooth size and long bone dimensions. Biostratigraphic and isotopic evidence indicates an age of ca. 54.5 Ma for the Cambay Shale vertebrate fauna, the oldest Cenozoic continental vertebrate assemblage from India, near or prior to the initial collision with Asia. Cambaytheriidae (also including Nakusia and Perissobune) and Anthracobunidae are sister taxa, constituting the clade Anthracobunia, which is sister to Perissodactyla. We unite them in a new higher taxon, Perissodactylamorpha. The antiquity and occurrence of Cambaytherium—the most primitive known perissodactylamorph—in India near or before its collision with Asia suggest that Perissodactyla evolved during the Paleocene on the Indian Plate or in peripheral areas of southern or southwestern Asia.
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RBINS Staff Publications 2020