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Article Reference Earliest perissodactyls reveal large-scale dispersals during the PETM
Perissodactyls were a diverse order of mammals in the Northern Hemisphere during the Paleogene, but very few species remain today. They first appear during the Paleocene–Eocene Thermal Maximum (PETM; 56 Ma) on the three continents of the Northern Hemisphere. Because they lack modern diversity, the study of their evolutionary history remains difficult, relying almost exclusively on fossils. In addition, their origin and early diversification remain elusive since all main groups appear at the same time with several genera and are already diverse, contrary to other modern mammal orders. Here, we investigate the early evolution of perissodactyls by analyzing a new dataset focusing on early species to obtain a new expansive phylogeny. This new topology, in combination with new observations and comparisons demonstrates that several genera are synonymous, simplifying the evolutionary picture of early perissodactyls. We show that the number of genera was overestimated and should be significantly reevaluated. These results also highlight fast dispersals of two genera, Pliolophus and Cardiolophus, on the three Northern continents, reflecting that of other modern mammals. The phylogeny also supports a potential Indo-Pakistani origin of perissodactyls. In addition, this large-scale phylogeny shows that many species that were named as “Hyracotherium” in the past (or sometimes “Eohippus”, the “dawn-horse”) and considered as “horses”, are not actually closely related to horses (Hippomorpha), including Hyracotherium itself, but can be considered as basal perissodactyls.
Located in Library / RBINS Staff Publications 2026 OA
Article Reference Early and High Medieval (c. 650 AD - 1250 AD) charcoal production and its impact on woodland composition in the Northwest-European lowland: a study of charcoal pit kilns from Sterrebeek (Central Belgium).
Located in Library / RBINS Staff Publications 2021
Article Reference Early and Middle Holocene human occupation of the Egyptian Eastern Desert: Sodmein Cave
Located in Library / RBINS Staff Publications
Article Reference Early Byzantine fish consumption and trade revealed by archaeoichthyology and isotopic analysis at Sagalassos, Turkey
We document the dietary and economic role of fish at Sagalassos, a town in ancient Pisidia (southwest Turkey) for the Early Byzantine period (c. 550 – 700 CE) through a detailed analysis of animal bones and stable isotopes. The role of fish in the diet is quantified, for the first time, based on large samples of sieved remains retrieved during the excavation of a number of spaces in an urban residence. The table and kitchen refuse from the mansion shows that fish was a regular part of the diet. However, past isotopic work focused on human individuals excavated in the city’s necropolises, slightly postdating the faunal remains examined, did not reflect this consumption of aquatic food. The studied assemblage comprises at least 12 different fish taxa, including five marine species, a Nilotic fish and six Anatolian freshwater species. Since the origin of the freshwater fishes could not be unambiguously determined by zoogeography alone, we analyzed carbon, nitrogen and sulphur stable isotope ratios in archaeological fish bones from Sagalassos as well as in bones of modern fish collected at different sites in Turkey. We show that most freshwater fish, i.e., all cyprinid species, came from Lake Eğirdir. No evidence was found for fish from the local Aksu River basin. The exact origin of pike, which account for 3% of all freshwater fish, could not be directly determined due to a shortage of modern comparative data. Using the data obtained on the provenance of the fish, the ancient trade routes possibly used in the Early Byzantine period are reconstructed using a combination of archaeological, numismatic and historical data on past commercial relations.
Located in Library / RBINS Staff Publications 2023
Article Reference Early cat taming in Egypt: a correction
A cat skeleton from a Predynastic burial in Egypt that was previously labelled as Felis silvestris is reidentified as Felis chaus. This means that the previous claim needs to be withdrawn that the specimen represents early evidence for taming of Felis silvestris that ultimately led to domestication. However, the statement that the small felid has been held in captivity for several weeks, based on the presence of healed fractures, is still valid.
Located in Library / RBINS Staff Publications
Article Reference Early Delphinida (Cetacea, Odontoceti) from the Miocene of the southern North Sea Basin
Located in Library / RBINS Staff Publications 2026
Article Reference Early diversification of seeds and seed-like structures
Located in Library / RBINS Staff Publications
Article Reference Early Eocene (Ypresian) continental vertebrate assemblage from India, with description of a new anthracobunid (Mammalia, Tethytheria)
Located in Library / RBINS Staff Publications
Article Reference Early Eocene artiodactyls (Mammalia) from western India
Located in Library / RBINS Staff Publications
Inproceedings Reference Early Eocene cambaytheres from Indo-Pakistan are the sister group of Perissodactyls
Cambaytherium, Nakusia, and Kalitherium are closely related early Eocene mammals from the Indo-Pakistan region that have been assigned to Perissodactyla (Laurasiatheria)or Anthracobunidae. The latter have been variously considered artiodactyls or perissodactyls, but more recently are usually placed at the base of the order Proboscidea or of the more inclusive Tethytheria (Afrotheria). We present new evidence from the dentition, skull, and postcranial skeleton of Cambaytherium, from Gujarat, India (ca. 54.5 Ma), that cambaytheres occupy a pivotal position as the sister taxon of Perissodactyla. Cambaytherium was more robust than basal perissodactyls such as ″Hyracotherium″ and Homogalax, and had a body mass of ~25-27 kg based on humeral, radial, and dental regressions. Perissodactyl synapomorphies include a transverse nasal-frontal suture, twinned molar metaconids, and an astragalus with deeply grooved trochlea and a saddleshaped navicular facet. Like perissodactyls, cambaytheres are mesaxonic and have hooflike unguals and a cursorially-adapted skeleton. Plesiomorphic traits compared to basal perissodactyls include bunodont molars with large conules and almost no hint of bilophodonty, unmolarized premolars, sacrum with four vertebrae, humerus with distally extensive pectoral crest and distal articulation lacking a capitular tail, distal radius without discrete scaphoid and lunate fossae, femur with low greater trochanter, calcaneus robust and wide with rounded ectal facet, astragalus wide with moderately long neck and vestigial astragalar foramen, navicular and cuboid short and wide, metapodials short and robust, and Mc I and Mt V present. In most or all of these traits cambaytheres are intermediate between phenacodontid condylarths and perissodactyls but closer to the latter. Our phylogenetic analyses place cambaytheres just outside perissodactyls, and place anthracobunids among primitive perissodactyls. However, similarities between cambaytheres and anthracobunids suggest that they are closely related, and future discovery of skeletal material of anthracobunids will provide a test of this hypothesis. Our results indicate that Anthracobunidae are not Proboscidea or tethytheres, and suggest that the origin of Perissodactyla may have taken place on the drifting Indian plate. How the progenitors of perissodactyls reached India is more problematic but might have involved land connections with Afro-Arabia during the Paleocene. Field work and research supported by the National Geographic Society.
Located in Library / RBINS Staff Publications