New material of the small raoellid artiodactyl Metkatius kashmiriensis is reported from the middle Eocene of the Upper Subathu Formation in the Kalakot area, Jammu and Kashmir, northwest Himalaya, India. The fossil material consists of numerous mandibular and maxillary fragments and isolated teeth, mainly belonging to juvenile specimens. It documents the poorly known dental morphology of M. kashmiriensis and provides an overview of its intraspecific variation, allowing to redefine its diagnosis. M. kashmiriensis is characterized by a particularly small size compared with other raoellid species, and by bunodont molars with moderately marked transverse lophs. The M/1–2 are much longer than wide and display characters similar to those of Rajouria gunnelli, such as the presence of a small paraconid and a mesial mesiostylid. The P/4 bears distally a small hypoconid, which appears to be unique in Raoellidae. The description of the new material also allows to document the poorly known morphology of the deciduous teeth of raoellids. The DP2/ is reported for the first time, and the DP/4 of M. kashmiriensis shows a morphology different from that of Indohyus, with the absence of mesial basin anterior to the paraconid and the primoconid. Contrary to what has recently been proposed, these results confirm that M. kashmiriensis is a valid species and not a synonym of Indohyus indirae, and highlight the great morphological diversity present within the Raoellidae during the middle Eocene in the Indian subcontinent.
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RBINS Staff Publications 2024 OA
tIn Becetèn, in southeast Niger, is one of the few mid-Upper Cretaceous (Coniacian-Santonian) fossil-iferous localities in Africa where multiple anuran taxa are known. Two of them, Pachycentrata taquetiand Inbecetenanura ragei, have been described from In Becetèn. Both are pipids, a clade composed ofexclusively aquatic anurans. Beside these two pipids, numerous isolated bones have also been referredto anurans within the site, but never referred to any known less inclusive taxa. Here we describe severalidentified cranial and postcranial bones referable to Pipidae or its stem-group, which together constitutePipimorpha. Among these elements, three cranial elements are referred to two unnamed pipimorphs.One of these has gondwanomorph affinities (pipids + South American and African extinct pipimorphs).These conclusions are supported by our phylogenetic analyses. Numerous postcranial elements are notreferable to any of the four pipimorph taxa identified at In Becetèn, although some ilia might be tenta-tively referred to ?Pachycentrata sp. The presence of at least four distinct anuran taxa makes In Becetènthe most diverse anuran site in Mesozoic Africa. It is also the first site in the Mesozoic where four pipi-morph taxa are identified. The dominance of pipimorphs in the anuran diversity of the site suggests thatIn Becetèn was likely more lacustrine than previously thought.
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RBINS Staff Publications 2025 OA
