Since the description of Isisfordia duncani, a number of new extinct species and revisions of previously described species have prompted a variety of contradicting phylogenetic hypotheses on the topology of Neosuchia. As a consequence, a consensus on the rooting of Eusuchia in relation to other neosuchian clades has not been reached and the origin of the group remains unsettled. Exemplifying this, Bernissartia fagesii, from the Early Cretaceous of Belgium, has long been considered a key taxon for understanding the origin of Eusuchia, but more recent hypotheses found support for a more basal position, as an ally to goniopholidids, paralligatorids or atoposaurids. Because many details of the anatomy of the type specimen are hidden by glue and the sediment adhering to the fossils, a number of characters are pending confirmation. Based on computed tomography data, we extract bones of the cranium and mandibles, describe new characters and re-evaluate anatomical details in the lectotype specimen. Our phylogenetic analysis confirms that B. fagesii is a derived neosuchian, unrelated to atoposaurids, goniopholidids and paralligatorids. We recover B. fagesii and Koumpiodontosuchus aprosdokiti in a basal position within Eusuchia, together with Susisuchidae, a group of gondwanan neosuchians containing Susisuchus and Isisfordia, which here form a polytomy with Hylaeochampsidae. The presence/absence of pterygoid-bound internal choanae cannot be used to fully resolve relationships at the neosuchian–eusuchian transition because of the variability of this character even at the familial level, as recently reported within susisuchids and bernissartiids. There is no doubt that true eusuchians were present in Laurasia as early as the Early Cretaceous, the hylaeochampsid Hylaeochampsa vectiana being the oldest (Barremian) undoubted representative. But whether the Eusuchia were also present in southern landmasses depends on solving the phylogenetic position of susisuchids and other less known gondwanan forms within or outside Eusuchia.
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Caverne X in Waulsort (Namur province, Belgium), excavated in the 19th century, revealed a burial site which was unexpectedly dated to the Final Upper Paleolithic (10,820 ± 80 BP, OxA-6856) in the 1990’s. A re-examination of the collection and a new radiocarbon dating program was recently undertaken. The dates obtained on four left femurs (9285 ± 30 BP, ETH-74725; 9310 ± 30 BP, ETH-74726; 9340 ± 30 BP, ETH-74727; 9300 ± 30 BP, ETH-74728) revealed that the remains should in fact be attributed to the Early Mesolithic,consistent with 24 other 14 C dates obtained for eight cave sites in the Meuse Basin which range from ca . 9600 BP to 9000 BP. Caverne X contained 544 human remains belonging to at least nine individuals (one fetus, one perinatal/young child, one teenager, two adolescents/young adults and four adults), and 66 faunal remains consisting mainly of intrusive animals with the possible exception of a cervid antler, and one artefact (a small flint blade). Other than ochre deposits, all alterations (breakage, surface abrasion, impact scars and concretions) are post-depositional in origin. Carbon and nitrogen stable isotope analysis indicates a diet primarily based on terrestrial resources from an open landscape with proteins provided by large herbivores. Our study shows that Caverne X fits well with results already obtained for the Meuse Basin cave burials in terms of chronology, minimum number of individuals, funerary rituals and diet.
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