Omomyiform primates are among the most basal fossil haplorhines, with the oldest classified in the genus Teilhardina and known contemporaneously from Asia, Europe, and North America during the PaleoceneeEocene Thermal Maximum (PETM) ~56 mya. Characterization of morphology in this genus has been limited by small sample sizes and fragmentary fossils. A new dental sample (n ¼ 163) of the North American species Teilhardina brandti from PETM strata of the Bighorn Basin, Wyoming, documents previously unknown morphology and variation, prompting the need for a systematic revision of the genus. The P4 of T. brandti expresses a range of variation that encompasses that of the recently named, slightly younger North American species ‘Teilhardina gingerichi,’ which is here synonymized with T. brandti. A new partial dentary preserving the alveoli for P1-2 demonstrates that T. brandti variably expresses an unreduced, centrally-located P1, and in this regard is similar to that of T. asiatica from China. This observation, coupled with further documentation of variability in P1 alveolar size, position, and presence in the European type species T. belgica, indicates that the original diagnosis of T. asiatica is insufficient at distinguishing this species from either T. belgica or T. brandti. Likewise, the basal omomyiform ‘Archicebus achilles’ requires revision to be distinguished from Teilhardina. Results from a phylogenetic analysis of 1890 characters scored for omomyiforms, adapiforms, and other euarchontan mammals produces a novel clade including T. magnoliana, T. brandti, T. asiatica, and T. belgica to the exclusion of two species previously referred to Teilhardina, which are here classified in a new genus (Bownomomys americanus and Bownomomys crassidens). While hypotheses of relationships and inferred biogeographic patterns among species of Teilhardina could change with the discovery of more complete fossils, the results of these analyses indicate a similar probability that the genus originated in either Asia or North America.
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RBINS Staff Publications 2019
The objective of the present revision is to complement the lithostratigraphy of the Ieper Group published in 2001 (Laga et al., 2001). This last publication reflected the activities in the Tertiary Subcommission at that time. The review published in 2001 framed in an initiative of the National Stratigraphic Commission and was limited to the lithostratigraphy at formation level. The Laga et al (2001) reference document has been the basis for the NCS website until now. The Ieper Group is characterised by clay−dominated sediments overlying, in most situations, the Landen Group strata and, if not outcropping, underlying the sand−dominated Zenne Group sediments. According to Laga et al. (2001) in their reference document for Paleogene and Neogene lithostratigraphy, the Ieper Group consists of the Kortrijk, Tielt and Gentbrugge Formations and ,members in these Formations are only listed. These subdivisions are also used on the 1:50 000 geological maps of Flanders, edited in the last decades of the 20th century. Especially the additional description of the members, and where possible, horizons, identified in the Formations, made the present review necessary and also modifications at the formation level itself arisen since 2001 needed to be integrated in a new synthesis. The present update is based on the earlier description of members in Maréchal & Laga (1988), Geets et al. (2000) and Steurbaut (1998) as far as appropriate. All modifications, discussions and additions are supported by the relevant literature references.
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RBINS Staff Publications 2016