We describe the abundant faunal remains that were found in an extensive ritual deposit discovered in 2012 at Oxyrhynchus. This site in Middle Egypt has been famous since the first millennium BC for the mormyrid fish that were worshipped there and after which the town was named. The role played by these fish has already been amply documented through textual evidence, bronze statuettes and paintings, but until now, no remains and no mummies of these fish had been found. We first describe the ritual deposit as a whole, with emphasis on its extent, its stratigraphy and its relationship to the surrounding structures, which, together with a very specific artefact, allow the layers to be dated to the Late Period. The fish remains, as well as the sparse mammal bones, are quantified using both number of identified specimens (NISP) and minimum number of individuals (MNI). Body length reconstructions of the mormyrid fish are carried out using newly derived regression equations. Because of the large quantity of material, we performed the taxonomic identifications and size reconstructions on subsamples from which estimates were then made for the total number of fish that may have been present in the entire deposit. Attention was given to the way in which the fish bundles were prepared, a process that involved both the use of textiles and halfa grass, and to how the deposit was organised. We discuss the species spectrum in relation to both the Egyptian fish cult and evidence from written sources. Finally, we attempt to reconstruct the different events that may have taken place between the capture of the fish and their final deposition at the site, using a combination of both zoological/ecological and papyrological evidence.
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1. A major goal in ecology is to unravel how species assemblages emerge and how they are structured across the landscape. Host–parasite systems are particularly interesting in this context, as limited host dispersal may promote the differentiation of parasite communities. 2. We examined whether the patterns of species diversity in Cichlidogyrus, a genus of monogenean parasitic flatworms with a direct life cycle, are consistent with the hypothesis that parasite diversity is driven by host dispersal. This was carried out by comparing two sympatric cichlid hosts (Tropheus moorii and Simochromis diagramma) with contrasting dispersal abilities. Genetic connectivity among host populations along the Zambian shoreline of Lake Tanganyika was estimated using microsatellite genotyping. Cichlidogyrus parasites were isolated and identified morphologically to the species level. 3.Simochromis diagramma, a host with a high dispersal capacity, was infected by a low number of Cichlidogyrus species, and the parasite assemblages were similar among host populations. In contrast, T. moorii, a host with a low dispersal capacity, was infected by a large number of Cichlidogyrus species, and the parasite assemblages differed strongly among host populations. These outcomes were thus as expected from the hypothesis. 4. Because of the strong host specificity of these Cichlidogyrus species, a lack of connectivity among host populations might facilitate allopatric speciation of the parasite.
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